Fistulina hepatica’s anamorphic version: Confistulina

Having been informed earlier by a friend that they have an article coming out soon in the journal Field Mycology and having then read the article, I considered it important to build on the information and photos shared in the manner I am most familiar: ramblings. I do not want to spoil the article so please do source the article yourself, though as a form of executive summary, this fungus, Confistulina hepatica, is the anamorphic (asexual) stage of the fungus Fistulina hepatica (beefsteak), which is common on oak and sweet chestnut. I have found it once on beech, though it generally sticks to the first two hosts. This anamorphic stage produces asexual spore, which adorn the fruiting body’s outer portion in abundance. The reason for its emergence is not known, though it might be weather-related. More information is absolutely required from across its host range – not just England!

Disclaimer: The older finds, because I didn’t know what I was looking at, are quite poor photographs. A huge shame, but alas!

Disclaimer 2: I suspect many of these are the anamorphic stage Confistulina hepatica, though none were confirmed so please do not assume they all are. This blog post is simply to begin building a knowledge base that builds of the limited resource pool of present.

The first oak that I’ll share is perhaps one of the more interesting of the bunch because, two years in a row (2015 and 2016), the outwardly anamorphic fruiting body appeared in the same location; albeit, in 2015, the fruiting body was larger. Nonetheless, it infers that, in a crude manner, the same mycelial colony has done the same thing two years in succession. Unfortunately, during 2015, I didn’t get a shot of the whole tree so the location of the arrow points to the 2016 occurrence. However, from the positioning of the ivy, we can see the location is the same.

Confistulina hepatica Fistulina anamorphic oak Quercus 1
The arrow points to the location, which is on the southern side of the tree right at the base.
Confistulina hepatica Fistulina anamorphic oak Quercus 2
The 2015 version in its rotten glory!
Confistulina hepatica Fistulina anamorphic oak Quercus 3
The fruiting body sports what looks like burns, in addition to exuding a tar-coloured liquid alongside the more routinely observed reddish exudations common to young fruiting bodies.
Confistulina hepatica Fistulina anamorphic oak Quercus 4
A closer look at the darker liquid from the 2015 version.
Confistulina hepatica Fistulina anamorphic oak Quercus 5
And a puncture wound (so it appears) in the fruiting body.
Confistulina hepatica Fistulina anamorphic oak Quercus 6
And the smaller 2016 emergence!
Confistulina hepatica Fistulina anamorphic oak Quercus 7
Again, we can see the darker liquid exudations. The morphology is also somewhat similar.
Confistulina hepatica Fistulina anamorphic oak Quercus 8
Closer still on the 2016 find.

The next series of shots were taken on my mobile phone back in 2015 and again from oak. Once more, the adornment of the fruiting body with exudations, of which some are darker, can be observed. 2016 saw the fungus return, with a vengeance, and in a different position, again, an anamorphic fruiting body.

Confistulina hepatica Fistulina anamorphic oak Quercus 9
Yes, this oak has been hammered. Poor tree! (2015)
Confistulina hepatica Fistulina anamorphic oak Quercus 10
At the base we can see the fruiting body.
Confistulina hepatica Fistulina anamorphic oak Quercus 11
Oozing everywhere!
Confistulina hepatica Fistulina anamorphic oak Quercus 12
Looks like something out of Alien, quite honestly!
Confistulina hepatica Fistulina anamorphic oak Quercus 34
2016 checking in. What a load of rubbish!
Confistulina hepatica Fistulina anamorphic oak Quercus 35
Further round the tree (note that the 2015 location was to the right) sits an anamorphic fruiting body.
Confistulina hepatica Fistulina anamorphic oak Quercus 36
A close(ish) look.
Confistulina hepatica Fistulina anamorphic oak Quercus 37
Closer still.
Confistulina hepatica Fistulina anamorphic oak Quercus 38
And closer yet further.

From here, we go to another 2015 find. This time, the fruiting body was a little elevated on the oak and evidently emanating from an area of burring / accumulation of dormant buds beneath the bark surface. This one quickly became senescent after perhaps a week so whether it’s Confistulina or not is tough to say – I include photos of both times I visited it. It did not reappear in 2016.

Confistulina hepatica Fistulina anamorphic oak Quercus 13
The first visit to this fungus, which was very small – maybe 5cm across.
Confistulina hepatica Fistulina anamorphic oak Quercus 14
A closer look.
Confistulina hepatica Fistulina anamorphic oak Quercus 15
A week later it had senesced.

The remainder of the photos take us into 2016 and most (but not all!) are taken with a better camera, which is good! First, we venture down to the New Forest and look at a well-decayed oak log, upon which a cluster of fruiting bodies sit atop the log.

Confistulina hepatica Fistulina anamorphic oak Quercus 16
Yes, yes, this photo was taken with a potato.
Confistulina hepatica Fistulina anamorphic oak Quercus 17
A very odd form but once again we can see the tarry liquid.
Confistulina hepatica Fistulina anamorphic oak Quercus 18
A side profile. Looks like a toe!
Confistulina hepatica Fistulina anamorphic oak Quercus 19
Yep, definitely a toe – a wrangled one, at that.

The next series are taken a month apart (late August and late September). What’s curious with this one is that we can see two anamorphic fruiting bodies and, come September, Laetiporus sulphureus fruiting directly alongside. The host, as can be seen, is oak, which is in a hedgerow of oak and ash.

Confistulina hepatica Fistulina anamorphic oak Quercus 20
Fading light caused the blur!
Confistulina hepatica Fistulina anamorphic oak Quercus 21
At the base we can see a dueo of fruiting bodies.
Confistulina hepatica Fistulina anamorphic oak Quercus 22
The first…
Confistulina hepatica Fistulina anamorphic oak Quercus 23
…and the gruesome second.
Confistulina hepatica Fistulina anamorphic oak Quercus 25
A month later a wild chicken appeared!
Confistulina hepatica Fistulina anamorphic oak Quercus 26
Two, in fact. Here we see the beefsteak and chicken side-by-side.
Confistulina hepatica Fistulina anamorphic oak Quercus 27
And here it’s in the distant murk.

Now, we move to a curious case of two fruiting bodies next to one another, again on oak, where one became a teleomorph and the other an anamorph. Why? Who knows. Very interesting, however, as I am sure you can appreciate.

Confistulina hepatica Fistulina anamorphic oak Quercus 28
An oak pollard by a swing. Look at the old pollard head.
Confistulina hepatica Fistulina anamorphic oak Quercus 29
A duo forming.
Confistulina hepatica Fistulina anamorphic oak Quercus 30
Looking pretty similar. Now watch…
Confistulina hepatica Fistulina anamorphic oak Quercus 31
…they change! No longer are they twins.
Confistulina hepatica Fistulina anamorphic oak Quercus 32
Exactly why this happened is a mystery though I’d really like to know – anamorph left) and teleomorph (right).
Confistulina hepatica Fistulina anamorphic oak Quercus 33
A side profile for comparison with the earlier one taken from the same position.

In October of 2016, I came across this majestic oak in a field. On a buttress root was what appeared to be an anamorphic fruiting body of Fistulina hepatica, which we can observe below.

Confistulina hepatica Fistulina anamorphic oak Quercus 39
Yup – majestic.
Confistulina hepatica Fistulina anamorphic oak Quercus 40
Also majestic??? (not sure!!)
Confistulina hepatica Fistulina anamorphic oak Quercus 41
Eh, nope. A bit more ugly, to be fair. Has the outward character of an anamorph though please don’t assume it is (wasn’t confirmed via microscopy).

Lastly, here’s one I suspect on sweet chestnut. I didn’t return to check how it did, though this year I’ll certainly keep more of an eye out and be more thorough in my inspections of the samples! Please share any examples you might have found of this, too – it’s important we build up a knowledge base.

Confistulina hepatica Fistulina anamorphic sweet chestnut Castanea 1
A lovely old coppice stool.
Confistulina hepatica Fistulina anamorphic sweet chestnut Castanea 2
Here sits the sample.
Confistulina hepatica Fistulina anamorphic sweet chestnut Castanea 3
Whether this is or is not is tough to say, though it does resemble an anamorphic stage somewhat.
Fistulina hepatica’s anamorphic version: Confistulina

David Attenborough on Richmond Park, London

Nothing much need be added, in light of who is narrating. As a 20-minute long film, it’s something you can readily watch at any point where you have some time going spare. There’s a few good segments on trees, including on ancient trees, deer, deadwood and wood-decay fungi. Really a fascinating watch!

Support the Friends of Richmond Park here.

David Attenborough on Richmond Park, London

Book review: Manual of Tree Statics and Tree Inspection

First and foremost, for those interested in purchasing the English version of the book, you can do so via the publisher Patzer Verlag, Summerfield Books or the Arboricultural Association. It’s not a cheap book and thus, for many, demands strong consideration before purchase, which is one of the core tenets behind why I wanted to read and review this book promptly following on from its publication. Therefore, my intention with this review is to give more information about the book over and above simply the listing of the chapters and sub-chapters, which are provided by the book sellers. Hence, interspersed amongst my review are photographs of the book, which will hopefully help guide you in your decision-making process. In fact, here’s an image for you, off the bat!

Quite a nicely presented hardback. The select illustrations you see on the front here occur throughout the book – hardly a page goes by where there isn’t at least one such image.

As regards the content, therefore, the book begins with a very succinct introduction followed by an extensive assessment of the countless aspects pertaining to tree biology. This biology section is adorned with illustrations that, as is the case across the entire book, help greatly with the simple conveyance of the core messages put forth by the authors – indeed, as is so prevalent and well-received across Mattheck’s works (in spite of the disagreements the autors have with the t/r ratio and h/d ratio, which they detail within). Somehow, I doubt this inclusion of illustrations is coincidental and instead a homage or emulation of what works well when describing detailed concepts in only a few words. Critically, with a book detailed as one that covers tree statics, the incorporation of tree biology into the statics model is included, which keeps the book on track from the beginnings and saves it from veering off course into just one that echoes prior understanding.

At this point, an interjection is necessary. Specifically, for the purpose of highlighting some of the very curious qualities of this chapter – in the good sense. Principally, an addition to the developing understanding of summer branch drop is provided, whereby the authors detail that, on top of the impact drought has on the failure of large branches, temperature plays a possibly significant role. Why, you ask? Because, as the authors detail, the pretensioning of a tree’s wood fibres (i.e. the manner in which it loads itself under its own weight and optimises itself for this) is lessened by heat, which relaxes the wood fibres – including atop branch junctions. When compiled with the fact that trees are much stronger under tension than compression, the developing compression zones underneath the branch junction (and just behind the branch collar) as the wood fibres relax atop build up and, alongside other factors (including drought and probably many other unknown variables), the branch can subsequently suddenly fail. The authors delve into this phenomenon more in the third chapter and beyond. Other intriguing and enlightening aspects of this second chapter include the remarks on how construction works impact upon tree stability, in addition to the efficacy of ivy to be both a bane and a blessing for the tree, as regards oscillation and damping during wind loads.

It is however the third chapter that arguably provides the most appreciable benefit to the reader, as it takes us into the realm of tree statics in the direct sense. The intricacies of the chapter will, of course, largely evade this review, as the intention of this post is not to ‘spoil’ the contents of the book to any marked degree and make its purchase redundant. Instead, what can be said is that it delves effectively into critical facets of statics and, in my personal opinion, the segment on wind loading and the impact of crown architecture and elevation on loading forces is most brilliant – even down to the illustrations and mathematical examples, which really do simplify an evidently complex engineering approach to tree risk assessment.

A segment from this third chapter, demonstrating how the authors blend text with diagrammatic models and more general illustrations.

Moreover, the discussion on the importance of hollowness for trees and why it should not always be considered a concern to the tree inspector is welcomed (unless it’s an open cavity and then, please, do be more worried straight off of the bat). In fact, and I quote: “the dimensions of a cavity in a tree have absolutely no informative value, as long as the occurring loads are not known.” Therefore, the t/r ratio Mattheck provided, the authors allege, is meaningless and incorrect.

Amazingly (from a factoid perspective), the authors then detail that a tree with a dbh of 1m and a central cavity leaving only 10cm of sound radial wood around the circumference has the same load-bearing capacity as a tree that is completely sound and has a dbh of 84cm! Of course, from an angle of understanding where the maximum loads occur, the fact that wood fibres stretch and compress far more at the outer sections of the stem than they do internally (with the neutral axis generally being the centre) gives credence and context to this assertion. The understanding of hollowness on older (veteran) trees is also discussed in a very articulate manner and, as it so seems, the greater risk for such trees is not failure of the trunk or root plate but of large lateral limbs attached to a hollowing trunk that can no longer sufficiently support such a mechanical load.

On this note, the outlining of root plate architecture is also deserving of mention. In this third chapter, the authors do a superb job of explaining why and how root plates fail and why, for all intents and purposes, extensive buttressing and adventitious rooting is not necessarily to be looked upon as defective. Using the example of a mature beech (see a below photo that I took recently), they allude to the often very pronounced root plate acting as a counter-balance to the lever arm that is the trunk (think of a very wide-based wine glass and compare it to a narrow-based one – which tilts more readily?). Indeed, where fungal decay is evident, as in the below photo, the effectiveness of the wide root plate comes into question, however.

Fagus sylvatica mature buttressing Ganoderma Meripilus 8
The admirable root plate of a mature beech. In this case, Ganoderma australe is compromising the butt and principal rooting structure, though where decay is not evident or is limited then a wider root plate is very effective at supporting the tree during times of wind loading – certainly more so than where such a wide root plate is not evident, on mature beech.

At this point, the notion of slenderness also enters the equation and, very curiously, the book asserts that slender multi-stemmed trees are more likely to oscillate excessively in wind loading conditions and fail, when compared to single-stemmed trees of equal slenderness. However, more crucially, why slenderness equals higher oscillatory frequency and thus denotes a greater risk of failure is detailed, which drives home the importance of not thinning out groups of trees and expecting the remaining ones to be able to stand wind-firm (the so-called ‘domino effect’ is also defined, whereby trees at a woodland edge successively fail, as those around become exposed after an initial tree fails and then those around it also fail).

And my favourite quote(s) of the third chapter? In reference to reaction growth laid down by the tree due to decay or other structural matters: “the development of symptoms is an expression of the vitality of the tree, not of its weakness” and that “focussing on so called symptoms can be absolutely counterproductive” – assuming, of course, you don’t quantify the weakness by ascertaining its load-bearing capacity (which is where statics comes in). Assumptions, therefore, are not good (though we all do assume, every day, when we assess trees, when working with outward symptoms solely).

A segment of the fourth chapter. This double page spread is certainly one of the rarer examples of where illustrations do not dress the paper in abundance.

Moving into the fourth chapter, we’re first greeted with a rather novel take on tree assessment: do not negate the emotional aspect of the act. To be precise, from this, I understand it as the authors inferring that the intuitive assessment of a tree, from afar, takes into account both our logical deduction of its form (based on the scientific understanding of what denotes good structural architecture in trees) and surrounds (targets, exposure, etc) and also its innate emotional ‘appeal’ (i.e. is the tree ‘harmonious’ and flowing well, or is it a rough and jagged structure that is abrasive to the eye and clearly there is something awry that demands more investigation). Here, the authors then throw in the concept of a female breast being harmonious (and not at all like a brick…!) to a child, who can appreciate the allure of that particular aspect of female anatomy for its biological purposes. I presume this wasn’t a translation error (of which there are a few, across the entire book, but it largely flows very well)! Thus, trees are like female breasts: harmonious (I could end the review here, quite frankly, for the comedic value).

Anyway, this fourth chapter then goes on to outline, to a very detailed degree, with the support of various diagrams in the sixth chapter (the ‘Annexes’), more about SIA (static integrated assessment). For something that I admittedly had considered quite complex, which it still certainly is, the authors do a stellar job in simplifying the concept and utilise diagrams and drawings to help ensure the reader can understand the message being conveyed. The explanation of the rationale behind why this methodology, provided in preceding chapters and also in this one, helps to contextualise the methodology and, quite truthfully, it’s a form of tree assessment that I am now keen to put into some form of practice. The statement the authors make about microdrills essentially being redundant now that sonic tomography has arisen from the mulch is also a curious one, which is deserving of some consideration, when appreciating that the former causes damage to trees and the latter, almost wholly, does not; in addition to the ability for a tomograph (i.e. PiCUS or ARBOTOM) to plot an entire cross-section and not just track the path of a single drill bitand provide information from that sole path.

A further useful segment within the fourth chapter is that of the pests and pathogens of the leaves and shoots of many common European tree species and genera. Indeed, the assertion that horse chestnut leaf miner is not solely an aesthetic issue is a welcome one, as for whatever concerning reason that understanding of it being only an aesthetic pest is still accepted by some in the industry. The section also has a really nice little bit of information on Massaria disease of plane, with a distinct Teutonic angle (as one would expect, from German authors).

A double page spread of the section of pests and diseases of the shoots and leaves of trees common to Europe.

The (‘technically’) last chapter, the fifth, draws upon the information provided within all prior chapters and provides the reader with guidance on how to, following the identification of a need to manage the tree (be it a newly-planted tree or veteran in a park or construction site), effectively enact a management regime that will be of benefit to the longevity of the tree in the landscape (this is why we manage many trees in ways that doesn’t see them get felled). Indeed, this chapter presents a lovely succinct look at the need to strongly consider management options, as in deciding upon a route there is then, in theory, no going back upon that decision (once scheduled work has been completed).

The encouragement by the authors to refrain from using a rigid support system, be that system a stake for a young tree or steel braces for old trees with slender co-dominant limbs, and instead utilise a flexible and dynamic system, is therefore well-received, as they present their argument in a sound and logical manner. Importantly, for bracing, the authors even go so far as to detail what angle a brace should be applied at and the load that it should be able to bear relative to the size of the parts being connected, which elevates this book to a level beyond that of others that, from memory, do not provide as much detail. Their comments on propping with A-frames and not upside-down V-frames are also welcomed and articulated so well with so few words. Again, illsurations really help instruct the reader and break the text apart, thereby making the reading more ‘bite-sized’.

limb propping tree lateral load a-frame
This duo of props fails to suppprt the limb against lateral forces, given their design. In lacking an A-framed design, these props are thus not optimally positioned to keep the limb of this sycamore from failing under a torsional load.

Finally, the appendices presented in chapter 6, which support many of the assertions made in the earlier chapters, arrive. There exist some fabulous tables showcasing the wood properties of different tree genera (cellular make up, strength, etc), the speed at which tree genera move water up the trunk in metres per hour, the compartmentalisation ability of different species and genera, and a comparison of various forms of tree assessment and their strengths / weaknesses. Also within these annexes is information on how hollow trees fail and a description of principal wood-decay fungi and their significance. The fungal section is limited in detail and species diversity (because it’s only an annex and thus cannot possibly be as detailed as a dedicated book on fungi) though, concerningly, the brittle cinder (Kretzschmaria deusta) is described as a white rot in a comparitive table (whilst being correctly marked as a soft rot in its descriptive text). Proofing of the text should have picked this up, as it is perhaps the most significant mistake in the book (all other issues identified are largely translation errors or typos, from what I could spot). The comments on Perenniporia fraxinea preferring robinia (Robinia pseudoacacia) over any other host tree in urban areas where soil is compacted are, nonetheless, very interesting. Anecdotally, I can understand and agree with this assertion.

So, my thoughts to summarise? This is a very useful book that serves not only as a primer for the statics integrated assessment model but also for tree visual assessments on the whole. The eagerness of the authors to promote analytical thinking of the reader was pervasive throughout and, in some respects, this book could essentially serve as a 101 for tree inspections and management to any tree inspector. Of course, the title alludes to this by referring to tree inspection, though I admit I was taken aback (in a positive sense) on how much breadth this book covered in so few words and so many images. For me, the mix works well and therefore the looming threat of a wall-of-text was evaded. Perhaps it is just me, though I can pick up influences from Shigo and Mattheck, in the use of few choice words to describe complex matters and to support such words with harmonious (see what I did there?) images.

If you have around £115 to spend and want a book that you can refer to routinely, this will be worth purchasing. Don’t think you’ll be getting an impossibly wordy book that delves into tree statics down to the minute level, however. That is reserved for other literature and, in my perspective, rightly so.

Book review: Manual of Tree Statics and Tree Inspection

Book scores; including on tree statics (!)

The library amasses further numbers with the below six books, which arrived during this month. All, quite obviously, pertain to trees and the landscape, and of notable interest will be the new tree statics book that was recently translated into English from German and published. Expect a detailed review of this book in the coming few days (currently around half way through reading it).

As always, links to where you can buy the books are given at the end of this post. Note that those ordered from Oxbow were acquired for a good third of their suggested retail value, as they were doing a clearance sale. Quite honestly, for anyone who likes anything to do with landscape history and ecology, bookmarking this particular publisher and distributor will yield some awesome results. For tree lovers, the imprint Windgather Press is probably one to keep a tabs on, as many of the books published involve trees.

new book purchases tree statics

To the left, we have a very interesting book on the mapping of Norfolk by William Faden and how the landscape has changed from then through to now. Tom Williamson, one of the authors, is a prominent authority on landscape history and thus the book does delve into the rural landscape of Norfolk and, to a degree, arboreal heritage. I picked it up for £10 (down from £30), and as of the time of this blog post it is still on sale and can be purchased here.

Along the top row working from the left, we first have the new book on tree statics and visual assessment, which, as I mentioned, will feature as a detailed review in a coming blog post. It’s certainly an expensive book so it is one to research before purchasing. By all means, await my review before deciding. This book I acquired for £115 from Summerfield Books at this link.

Adjacent is what looks to be a truly captivating book on the Cedar of Lebanon and its prominence in human history. From my readings on this tree in the culture and religion of many historic peoples, this will be next on my list of reading material and will undoubtedly be rich in information that helps build my view of the tree further. As a very recently publication, it can be purchased via Archaeopress for £36 here. Please, consider trawling their book store as there are some other very great books they sell that involve trees and the natural landscape.

The lower duo are books on the historic gardens of Derbysire (left) and hedgerows across Britain (right). Following on from the information provided in Mark Johnston’s Trees in Towns and Cities: A History of British Urban Arboriculture, I have been keen to learn more about pleasure gardens and earlier arboretums / plant collections, so this book specific to Derbyshire will hopefully supplement my learning on this front and can be bought here from Oxbow Books. As for the latter, the detailed exploration of hedgerow characteristics and distribution throughout Britain is somthing that, would you have guessed, I also find intriguing. Indeed, hedgerows have historically – and still are – being gutted, for a variety of reasons, which means understanding what makes them unique (even down to a specific region) and crucial ecological and cultural features very necessary. This book is currently reduced from £25 to £10 on Oxbow Books and can be purchased here. The book is clad with images and can easily be read entirely in a day.

The last book, which is all the way to the right, further expands upon my desire to learn more about botanic gardens. Instead of Derbyshire however, this book takes the reader to Manchester – specifically, the Manchester Botanic Garden. At £5, though reduced from £25, its acquisition was a no-brainer and, from scanning the book, it appears an utter steal. Again, this was purchased via Oxbow Books via this link.

Keep an eye out for my review of the tree statics book. Hopefully, if all goes to plan, it’ll be online before the close of the bank holiday.

Book scores; including on tree statics (!)

Interesting cases of wood-decay fungi

I have been absurdly busy so haven’t been blessed with the time to get some blogging in, though I have been graced with thirty minutes of time this evening so without any further utterings we’ll delve right into the good stuff – trees and fungi (in my usual frenetic and incoherent manner). Plus, listening to some early Hawkwind has really got me in the mood to do something useful!

The cases are all from an absolutely splendid park down in Maidstone – Mote Park. Honestly, if you live anywhere near there, do pay it a visit and explore as much as you can (it’s massive!). The sheer abundance of mature and veteran trees provides for a magnificent display of fungi and, so I am told, there is a need to record the saproxylic insects on site on the many monoliths and moribund trees.

To kick this post off, I take you to a very interesting case of Pseudoinonotus dryadeus on oak – three of them, all of which are within 8-10m of one another and share a crown. All fruitings of the fungus are historic though its presence on all three trees makes for some tempting considerations – namely, the synchronicity of fruiting (are they similar genotypes?) and the means of colonisation (spore or something else?). Indeed, I can only infer some sort of fungal mysticism or sorcery in positing both aspects for consideration (there is no proof of either, per se), though it did make me think. Perhaps it will make you readers think as well! (!?)

Pseudoinonotus dryadeus colonisation senescent old 1
The recipients of the Manchurian Candidate!
Pseudoinonotus dryadeus colonisation senescent old 2
Exhibit one!
Pseudoinonotus dryadeus colonisation senescent old 3
At the base (to the left).
Pseudoinonotus dryadeus colonisation senescent old 4
Quite an old bracket but a bracket nonetheless. Lovely buttressing, too.
Pseudoinonotus dryadeus colonisation senescent old 5
Exhibit two. More brackets and more buttressing.
Pseudoinonotus dryadeus colonisation senescent old 6
Shame these got yanked off, too.
Pseudoinonotus dryadeus colonisation senescent old 7
Exhibit three! SOme glorious buttressing here, yet again. Thus the fungus gets its common name: the Eiffel Tower fungus.
Pseudoinonotus dryadeus colonisation senescent old 8
Old, dead (not really but who cares for semantecs?) but not forgotten.

And then…and then…more Pseudoinonotus dryadeus – literally 100 yards down the same path. Oh how Mote Park delivers! This example also really does demonstrate the magnificent buttressing induced by its decay on oak, as you’ll see.

Pseudoinonotus dryadeus mature oak butt decay 1Pseudoinonotus dryadeus mature oak butt decay 2Pseudoinonotus dryadeus mature oak butt decay 3Pseudoinonotus dryadeus mature oak butt decay 4Pseudoinonotus dryadeus mature oak butt decay 5

Would you then believe it? Essentially opposite (no joke) were two colossal beech trees fenced-off (as if that ever stopped me??!) that, as anyone who has seen mature or veteran beech buttressing all over the place like egg whites pour out of a broken egg when broken too aggressively (nice analogy? – likely not), drew me in. Was I disappointed? Not at all! Ganoderma australe and Meripilus giganteus all over the option.

Fagus sylvatica mature buttressing Ganoderma Meripilus 1
Good cop (right) bad cop (left) – something something pun something something copper beech and tell better jokes
Fagus sylvatica mature buttressing Ganoderma Meripilus 2
The copper beech to the right with roots all over the option.
Fagus sylvatica mature buttressing Ganoderma Meripilus 3
Ganoderma australe and Meripilus giganteus – dual decay. Decay squared? That raises a good thought – is decay by more than one fungus simply a cumulative issue or instead a geometric or even negatory issue (i.e. is decay of two types ‘less serious’ than from one only)? Question galore and no answer. Someone ask an expert!
Fagus sylvatica mature buttressing Ganoderma Meripilus 4
Merip (foreground) and Gano (background). Also plenty of blades of grass for the monocot enthusiasts who stumbled across this blog because I wrote the word monocot.
Fagus sylvatica mature buttressing Ganoderma Meripilus 5
Ganodeerma australe being illuminated by a sunburst.
Fagus sylvatica mature buttressing Ganoderma Meripilus 6
The diddy little brackets further up the stem weren’t blessed with sun.
Fagus sylvatica mature buttressing Ganoderma Meripilus 7
And between some more buttresses there were some more Ganodermas.
Fagus sylvatica mature buttressing Ganoderma Meripilus 8
Onto the plain old beech now. Exquisite buttressing here!
Fagus sylvatica mature buttressing Ganoderma Meripilus 9
Did someone say fungi?
Fagus sylvatica mature buttressing Ganoderma Meripilus 10
Nope – nothing to see here.
Fagus sylvatica mature buttressing Ganoderma Meripilus 11
Nor is there anything to see here. I’m just tired of writing captions!
Fagus sylvatica mature buttressing Ganoderma Meripilus 12
Oh look, finally. Something. Some nice husks. Probably some Ascomycetes on them (Xylaria carpophila).

To finish up, because I’m getting tired and I am up early tomorrow, here’s something to sit on whilst you ponder the plethora of ultimate questions spewed forth from my mind with little restraint – a dryad saddle. The host? Not sure – lots of ash about though one can never rule out sycamore (unless you’re in the middle of a Douglas fir plantation?). These had actually already over-matured, which means you can see dryad saddle (i.e Cerioporus squamosus – named, prior to that, Polyporus squamosus) out there if you look!

Dryad saddle Polyporus Cerioporus squamosus 1Dryad saddle Polyporus Cerioporus squamosus 2Dryad saddle Polyporus Cerioporus squamosus 3Dryad saddle Polyporus Cerioporus squamosus 4

Interesting cases of wood-decay fungi

Trees in the conflict of Israel and Palestine

Everything written here is supported by sources I have referenced (check for yourself), as always, so do not treat this as an assault on either side and / or their respective religions. This post is through the lens of the tree, so treat it as such. Moreover, the entire topic is very interesting.

As Israel collides with Palestine, trees are – and always have – been caught up in the melee. Principally, olive and citrus groves, some of which may have been tended to for many centuries by the Palestinians (Temper, 2009), are bulldozed or otherwise uprooted, with little respect for their cultural and historical importance (Allen, 2008; Graham, 2002). As an example, in 1986, when the Israeli military seized Midya, over 3,300 olive trees were uprooted, and a further 2,000 olives were bulldozed in Qattana (Bardenstein, 1999). Some of the trees removed from Qattana were later re-planted within the Jewish sector of West Jerusalem (Lentin, 2000), though by that point the damage (in many an aspect) had certainly been done. Some Israeli residents did protest their planting (out of anger towards the state), by tying ribbons to the trees that contained messages such as “Take me back to Qattana!” (Bardenstein, 1999), whilst others, across the entire conflict, have chained themselves to the olive trees in order to stop the bulldozers from uprooting them (Sfard, 2009), supported Palestinian farmers by helping them harvest from their olive trees and, at times, defending them in the process (Stephan, 2003), provided replacement olive tree for those uprooted (usually by settlers), or helped to retain olive trees within occupied territories for their symbolic meaning of peace (Braverman, 2009) – “extending the olive branch“, per se.

AN olive grove being bulldozed. Source: The Independent.

Below the surface level of removing ‘enemy’ trees, the removal of olive trees has a very political undertone. Olive trees have been held in very high regard by Palestinians for generations (and are regarded by some as holy trees), where they were farmed and thus supported viable economies (Braverman, 2009; Cohen, 1993), and their removal (or ‘capture’, by where groves were encompassed into the territory of Israel) by Israeli forces therefore can also be interpreted as an attack on Palestinian culture and custom (Bardenstein, 1999; Bowman, 2007; Braverman, 2009; Kershner, 2005) – notably when such acts are supported by the Court (Sfard, 2009). In some cases, it may even be Jewish settlers who vandalise or cut down the olive trees (Kershner, 2005), and even when the Israeli army have allowed the Palestinians to harvest their olive crops. In such instances, the Israeli army will generally not intervene (Pigni, 2010).

Such a political (and, to a marked degree, religious) act may be most pertinently discerned when the olive groves (or individual trees) are captured or destroyed during harvesting season, which has indeed occurred in some instances (Batniji et al., 2009). Moreover, the fact that many olive groves have been uprooted (comprising of tens of thousands of individual olive trees – in Qafeen alone, 12,600 olives were uprooted for this reason) for the construction of the Separation Barrier in the West Bank was also a cause of huge upset, for the Palestinians; particularly when their uprooting was coupled with justifications including to construct watchtowers, roads, checkpoints, and other security fences (all of which further hamper daily life and privacy), in addition to the use of the groves for sheltering armed Palestinians (Braverman, 2009). For those groves not uprooted, the Separation Barrier may instead have isolated Palestinian farmers from their olive trees, for much of the year. In Qafeen, over 100,000 olive trees suffered this isolated fate.

Olive trees are removed to facilitate the construction of the Separation Barrier. Source: Haaretz.

Whilst the capture and removal of Palestinian groves has been ongoing, Israel has also been afforesting barren regions of its territory – and for many decades. Spearheaded largely by the Jewish National Fund that was established in 1901 (and since 1961 has been Israel’s exclusive forestry agency), the afforestation program was, at its core, a religious, ecological, and territorial pursuit (Amir & Rechtman, 2006; Bardenstein, 1999; Braverman, 2009; De-Shalit, 1995; Ginsberg, 2000; Stemple, 1998; Tal, 2013), with pine species (including Pinus halepensis) being particular favourites (Osem et al., 2008; Weinstein-Evron & Galili, 1985). In recent decades, the emergence of numerous pests associated with the pine (such as Matsucoccus josephi) has however led to more diverse plantations, with other pine species (including Pinus brutia) and deciduous tree species being selected for use (Braverman, 2009).

In essence, a core reason for this afforestation is because Israel, in the ages gone by, was considered to be covered with forests (even up to the 11th century A.D., in places), though it is suggested that when the Jewish people were in exile those who occupied Israel (from around 722 B.C. – 1948 A.D.) destroyed many of these forests (due to arson, harvesting for fuel, overgrazing, sabotage, and warfare) and thus, upon the return of the Jewish people to Israel, in order to bring Israel back to its former character, forests were (and still are) planted upon the barren slopes (Stemple, 1998; Tal, 2012; Tal, 2013). Braverman (2009) states that the Jewish National Fund has planted over 200,000,000 trees across more than 225,000 acres of claimed land, since its inception. However, according to the Old Testament, in the book of Joshua, even Jewish peoples have been responsible for some of this historic clearance in their Promised Land (Tal, 2013), and for this reason the Jewish National Fund is seeking to restore Israel’s forests of ten thousand years ago – soon after the last glacial ice age. In fact, a great deal of planting, each year, is undertaken in the leading up to – and on the day of – Tu B’shvat (Bardstein, 1999; Zerubavel, 2000).

israel afforestation time contrast
Hiran Forest in 1998 (left) and 2008 (right). Source: KKL-JNF.

After the creation of the Jewish National Fund, though prior to its major afforestation practices towards the middle of the century, the British had, since 1918 (after they had seized southern Palestine), planted up many hundreds of thousands (if not many millions) of saplings (comprising of species including stone pine, tamarisk, terebinth, and oaks) on the hills of Israel (Tal, 2013), and before this (from 1860 onwards) the Turkish Ottoman Empire and settling German Templars had done much the same (Ginsberg, 2006; Liphschitz & Biger, 2004).

In this afforestation project, such planted areas are also oft designated as forest reserves and thereby protected by Israeli law, which Braverman (2009) dubs as “lawfare” against the Palestinians, whose land may have been afforested following seizure. This planting up of forest on occupied lands, of which a sizeable portion was planted over destroyed Palestinian villages in the years after 1948 (an act of camouflage, and for some allegedly the camouflage of war crimes), also makes the land very difficult to reclaim, as the reclaimers must first remove all of the trees (after gaining the permission to clear the perhaps protected forest); in this sense, Palestinians may never be able to occupy such land again, be it for living within or for cultivation. In some cases, Braverman (2009) writes, Palestinians have even retaliated against this afforestation by firing rockets into the planted pine forests or burning the pine forests through arson, with a desire much aligned to Israel’s uprooting of the olive trees (in a sense, a ‘tree for a tree’). In this respect the tree, and specifically the pine, is a tool of war, and thus represents the enemy as a solider would (Boerner, 2011; Braverman, 2008).

On a more philosophical level, the fact that the Jewish National Fund would plant a tree for each newborn from Jerusalem in Jerusalem’s artificially-borne Peace Forest, dedicate the specific tree to the child, and provide the individual with a certificate (including a photo of the tree) that remarks on how it is hoped the tree and child grow together, outlines the innate affinity (or interchangeability) man has with trees (Braverman, 2009); as is detailed before this blog post on earlier ones associated with trees and religion.

Furthermore, the populist and globally crowd-funded nature of a fair portion of the tree planting, supported via financial gifts (complete with material rewards, such as memorial stones) and the use of the ‘Blue Box’ (located in households, schools, and offices), sewed into the fabric of the afforestation project a very emotionally evocative and inclusive aspect to both children and adults of the Jewish faith, even if the donator was geographically separated from Israel (Bar-Gal, 2003; Braverman, 2009; Zerubavel, 2000). Perhaps, this ability for a Jewish person to fund the planting of a tree may dampen their feeling of loss for not living within the Promised Land; in place of their presence, they can fund the planting of a tree, which can be considered a “proxy immigrant” (Braverman, 2009). At a tangent, the returning of the landscape to forest is also important on a cultural level, because the forests were incredibly important for the Jewish peoples’ ancestors; often would children be named after trees, and even Israel itself was sometimes compared to a tree (Zerubavel, 2000; Zerubavel, 2005).


Allen, L. (2008) Getting by the occupation: How violence became normal during the Second Palestinian Intifada. Cultural Anthropology. 23 (3). p453-487.

Amir, S. & Rechtman, O. (2006) The development of forest policy in Israel in the 20th century: implications for the future. Forest Policy and Economics. 8 (1). p35-51.

Bar-Gal, Y. (2003) Propaganda and Zionist Education: The Jewish National Fund, 1924-1947. USA: University of Rochester Press.

Bardenstein, C. (1999) Trees, forests, and the shaping of Palestinian and Israeli collective memory. In Bal, M., Crewe, J., & Spitzer, L. (eds.) Acts of Memory: Cultural Recall in the Present. USA: University Press of New England.

Batniji, R., Rabaia, Y., Nguyen–Gillham, V., Giacaman, R., Sarraj, E., Punamaki, R., Saab, H., & Boyce, W. (2009) Health as human security in the occupied Palestinian territory. The Lancet. 373 (9669). p1133-1143.

Boerner, R. (2011) Trees as soldiers in a landscape war. Landscape Ecology. 26 (6). p893-894.

Bowman, G. (2007) Israel’s wall and the logic of encystation: Sovereign exception or wild sovereignty?. Focaal. 50 (1). p127-135.

Braverman, I. (2008) “The Tree Is the Enemy Soldier”: A Sociolegal Making of War Landscapes in the Occupied West Bank. Law & Society Review. 42 (3). p449-482.

Braverman, I. (2009) Planted Flags: Trees, Land, and Law in Israel/Palestine. USA: Cambridge University Press.

Cohen, S. (1993) The politics of planting: Israeli-Palestinian competition for control of land in the Jerusalem periphery. USA: University of Chicago Press.

De‐Shalit, A. (1995) From the political to the objective: the dialectics of Zionism and the environment. Environmental Politics. 4 (1). p70-87.

Ginsberg, P. (2000) Afforestation in Israel: a source of social goods and services. Journal of Forestry. 98 (3). p32-36.

Ginsberg, P. (2006) Restoring biodiversity to pine afforestations in Israel. Journal for Nature Conservation. 14 (3). p207-216.

Graham, S. (2002) Bulldozers and bombs: the latest Palestinian–Israeli conflict as asymmetric urbicide. Antipode. 34 (4). p642-649.

Kershner, I. (2005) Barrier: the seam of the Israeli-Palestinian conflict. USA: Palgrave Macmillan.

Lentin, R. (2000) Israel and the Daughters of the Shoah: Reoccupying the Territories of Silence. USA: Berghahn Books.

Liphschitz, N. & Biger, G. (2004) Green Dress for a Country – Afforestation in Eretz Israel: The first hundred years 1850-1950. Israel: KKL.

Osem, Y., Ginsberg, P., Tauber, I., Atzmon, N., & Perevolotsky, A. (2008) Sustainable management of Mediterranean planted coniferous forests: an Israeli definition. Journal of Forestry. 106 (1). p38-46.

Pigni, A. (2010) A first-person account of using mindfulness as a therapeutic tool in the Palestinian Territories. Journal of Child and Family Studies. 19 (2). p152-156.

Sfard, M. (2009) The Price of Internal Legal Opposition to Human Rights Abuses. Journal of Human Rights Practice. 1 (1). p37-50.

Stemple, J. (1998) Viewpoint: a brief review of afforestation efforts in Israel. Rangelands. 20 (2). p15-18.

Stephan, M. (2003) People power in the Holy Land: How popular nonviolent struggle can transform the Israeli-Palestinian conflict. Journal of Public and International Affairs. 14 (Spring). p164-183.

Tal, A. (2012) Israel’s New Bible of Forestry and the Pursuit of Sustainable Dryland Afforestation. Geography Research Forum. 32 (1). p149-167.

Tal, A. (2013) All the Trees of the Forest: Israel’s Woodlands from the Bible to the Present. USA: Yale University Press.

Temper, L. (2009) Creating facts on the ground: Agriculture in Israel and Palestine (1882-2000). Historia Agraria. 48 (1). p75-110.

Weinstein-Evron, M. & Galili, E. (1985) Prehistory and paleoenvironments of submerged sites along the Carmel coast of Israel. Paleorient. 11 (1). p37-52.

Zerubavel, Y. (2000) The Forests as a National Icon: Literature, Politics, and the Archeology of Memory. In Elon, A., Hyman, N., & Waskow, A. (eds.) Trees, Earth, and Torah: A Tu B’Shvat Anthology. USA: The Jewish Publication Society.

Zerubavel, Y. (2005) The forest as a national icon: literature, politics, and the archaeology of memory. Israel Studies. 1 (1). p60-99.

Trees in the conflict of Israel and Palestine

Street Trees in Britain: A History (new book, October 2017)

As I was trawling the depths of the Oxbow Books store, which is an utter treasure trove of books on various matters (many pertain to trees, hedgerows, gardens, woodlands, forests and the broader concept of landscape ecology – notably from the imprint Windgather Press), I came across a new book, to be released later this year, by Mark Johnston, the author of Trees in Towns and Cities: A History of British Urban Arboriculture.

So what is the title of this book? If you paid attention to the title, you’d already know, though for the sake of clarification it’s Street Trees in Britain: A History. As regards a description of the text, taken from the publisher’s page:

The trees which line many of the streets in our towns and cities can often be regarded as part of a heritage landscape. Despite the difficult conditions of an urban environment, these trees may live for 100 years or more and represent ‘living history’ in the midst of our modern streetscapes. This is the first book on the history of Britain’s street trees and it gives a highly readable, authoritative and often amusing account of their story, from the tree-lined promenades of the seventeenth century to the majestic boulevards that grace some of our modern city centres.

The impact of the Victorian street tree movement is examined, not only in the major cities but also in the rapidly developing suburbs that continued to expand through the twentieth century. There are fascinating descriptions of how street trees have helped to improve urban conditions in spa towns and seaside resorts and also in visionary initiatives such as the model villages, garden cities, garden suburbs and new towns.

While much of the book focuses on the social and cultural history of our street trees, the last three chapters look at the practicalities of how these trees have been engineered into concrete landscapes. This includes the many threats to street trees over the years, such as pollution, conflict with urban infrastructure, pests and diseases and what is probably the greatest threat in recent times – the dramatic growth in car ownership.

Street Trees in Britain will have particular appeal to those interested in heritage landscapes, urban history and the natural and built environment. Some of its themes were introduced in the author’s previous work, the widely acclaimed Trees in Towns and Cities: A History of British Urban Arboriculture.

Retailing at £30, you can currently pre-order it for £22.50, which seems like a relative steal. I would strongly recommend that, if you are to pre-order it, you do so via the publisher here, which supports them directly and will enable them to continue to press more books of a similar nature. At the very least, they’ve got a pre-order through me, though given many who frequent this blog are from the United Kingdom, I hope they’ll receive more interest (I’m sure they will!) prior to the book’s publication.

Street Trees in Britain: A History (new book, October 2017)